"After designing and installing a very high end hand selected custom milled rustic oak floor..." "Do I have to install air conditioning?"
Seems like these two things are conflicting - you've put a lot of care and attention and probably money into this custom flooring project, obviously you've done your research into finishes and maintenance, yet you didn't plan for humidity control? One of the first conversations I have with any client, especially during a reno, is about climate control - the wood needs to be installed in the climate it is going to perform in, and that should be consistent all year round if you want the floor to behave consistently. This means monitoring your Relative Humidity all year round and using humidifiers or dehumidifiers as necessary. The goal range is 40-50%RH for most products and when you say humidity was in the 75-80 I really hope you mean outside and not inside the house! You'd be looking at buckling etc, all kinds of expansion related issues.
Short answer, YES get the AC going, that will help the circulation and may help your other issues. Most importantly however will be the dehumidifier - an AC unit will remove some moisture through condensation but not always enough depending on the house.
With regards to the finish - once you get your climate sorted, see if its still an issue - I know skylights and other concentrated areas where sunlight shines directly on the floor for long periods will lead to oxidization of fading/lightening of the stain over time. Sometimes a rug is the best/easiest solution.
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